The study is the first of its kind to explore if savanna-mosaic habitats would account for increased time spent on the ground by the Issa chimpanzees, and compares their behaviour to other studies on their solely forest-dwelling cousins in other parts of Africa. Overall, the study found that the Issa chimpanzees spent as much time in the trees as other chimpanzees living in dense forests, despite their more open habitat, and were not more terrestrial (land-based) as expected. Furthermore, although the researchers expected the Issa chimpanzees to walk upright more in open savanna vegetation, where they cannot easily travel via the tree canopy, more than 85% of occurrences of bipedalism took place in the trees.